Boris the Animal, last of the malevolent Boglodites that consumed all planets in their path, escapes from a lunar prison seeking vengeance on Men in Black Agent K, who shot off his left arm and jailed him in 1969. Boris confronts K and his partner Agent J, telling K he is "already dead" then fleeing. J searches the MIB database when K refuses to discuss his past, and discovers Boris committed several murders in 1969 before being captured by K, who then deployed an "ArcNet" shield around Earth that prevented a Boglodite invasion and starved the race to extinction. Agent O, the new head of MIB after Agent Zed's passing, tells J not to probe deeper. That night as K speaks with J on the phone, K vanishes. When J arrives at MIB headquarters looking for him, only he remembers K ever existed. O shows J a memorial statue of K, who was killed by Boris in 1969 — though she also notes that the chocolate-milk cravings and headaches J is experiencing are a result of a "temporal fracture" of time travel. As the Boglodites invade Earth, J learns Boris time-jumped to July 16, 1969, killed K, and prevented the ArcNet from being installed. J time-travels to the day before to try to save K.
In 1969, at Coney Island where the first murder took place, J finds 1969 Boris and attempts to shoot him but is knocked out by 1969 K. J is taken to 1969 MIB headquarters for questioning, and finds K flirting with 1969 O. K prepares to neuralyze J until J reveals his mission, convincing K to accept his aid in stopping Boris. They follow clues to Griffin, a fifth-dimensional being who sees multiple timelines simultaneously and can give them the ArcNet. At The Factory studio party hosted by Andy Warhol (who is revealed to be an MIB agent), 1969 Boris attacks. Griffin flees, after promising a gift and a miracle game at Shea Stadium. J remembers the "miracle game" of the 1969 World Series, and the agents arrive at Shea to find Griffin, who shares his timeline vision of "his favorite moment in human history", when the underdog New York Mets win the World Series. Before Griffin can give them the ArcNet, 1969 Boris kidnaps him. J and K pursue and rescue Griffin and the ArcNet, but 1969 Boris escapes and meets his 2012 counterpart.
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